Day 2 companies make high-quality decisions, but they make high-quality decisions slowly. To keep the energy and dynamism of Day 1, you have to somehow make high-quality, high-velocity decisions.
There is no such thing as great wisdom in the world, just as there is no such thing as talent. What exists is triviality. If you keep doing repetitive things, you become an expert; if you do repetitive things with special focus, you become a master.